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The film deals with the situation of British prisoners of war during World War II who are ordered to build a bridge to accommodate the Burma-Siam railway. Their instinct is to sabotage the bridge but, under the leadership of Colonel Nicholson (Alec Guinness), they are persuaded that the bridge should be constructed as a symbol of British morale, spirit and dignity in adverse circumstances. At first, the prisoners admire Nicholson when he bravely endures torture rather than compromise his principles for the benefit of the Japanese commandant Saito (Sessue Hayakawa). He is an honorable but arrogant man, who is slowly revealed to be a deluded obsessive. He convinces himself that the bridge is a monument to British character, but actually is a monument to himself, and his insistence on its construction becomes a subtle form of collaboration with the enemy. Unknown to him, the Allies have sent a mission into the jungle, led by Warden (Jack Hawkins) and an American, Shears (William Holden), to blow up the bridge.

1943. With the battle of Stalingrad turning the war against them, the Germans are attempting to bully neutral Turkey into joining the Axis; to this end they have trapped 2,000 British soldiers on Kiros, an island in the Aegean, with only one sea route for evacuation, a sea route commanded by two gigantic German antiship batteries deployed in a massive cliffside bunker on the island of Navarone. Immune to air attack and too much for Allied battleships to suppress, the British muster Keith Mallory, a commando officer who has been working on occupied Crete for nearly two years and who is an expert mountaineer, to ferry a team of British commandos to the only area of Navarone that is not monitored by the Germans, a 400-foot cliff. Greek resistance is to meet the team inland and guide them around German patrols to the area of the German guns. However, the commanding officer of the British team suffers grave injury in the climb and Mallory must take control of the mission, despite clashes with explosives expert John Anthony Miller, who upon the arrival of the night of the raid finds his equipment has been sabotaged, thus exposing a traitor in the team’s ranks.